The game's protagonist, (based on war hero William Grover-Williams), is an Irish man named Sean Devlin, a regular among the racing groups of Paris. During a race in the Saarbrücken against Kurt Dierker, a prominent Nazi, Kurt shoots out Sean's tires when it becomes apparent that Sean will win. Sean and his best friend named Jules follow Kurt and steal his car before driving it off a cliff. Unfortunately for the pair Dierker is a Nazi secret agent and the two are swiftly captured and interrogated by Dierker at the Doppelsieg factory. Dierker is convinced they are involved with British Military Intelligence and executes Jules when he and Sean deny this. Sean manages to escape and returns back to Vittore's farm, only to find it being burned to the ground. After rescuing Jules' family, Vittore and Veronique. the three escape to the Cabaret in Paris, belonging to Vittore and Veronique's Family; the Belle De Nuit.
Sean is rebuked and recruited by Luc Gaudin, a French Resistance fighter. During his escapades, Sean is contacted by Skylar, a femme fatale British racer and Sean's old flame. Skylar drugs Sean before presenting him to Bishop and associate Wilcox,who reveals that Skylar is a British Intelligence officer and recruits Sean, in exchange he promises to aid Sean in hunting down and killing Dierker. Sean agrees and following Bishop's orders sneaks into a mausoleum retrieve a mysterious chest, which the Nazis have obtained. After returning the chest to Bishop and Wilcox, Bishop tells Sean that Dierker will be leaving for Berlin in a Zeppelin currently docked at where Sean is to deliver the chest to. Sean eventually manages, through the help of Skylar, to enter the Zeppelin, where he encounters Dierker alone. In a fierce struggle between the two men a stray gunshot from Dierker's pistol ignites the hydrogen gas cells of the Zeppelin.Dierker escapes with a parachute ,leaving Sean with no other choice but to jump from the zeppelin before it explodes.
Luc upon Sean's return, tells Sean to contact another Resistance cell and gives him the name of a contact who can provide the papers needed to bypass checkpoints. Sean meets up with the a man named Santos, who supplies Sean with the transit papers. Sean meets up with Margot, the leader of the French Resistance in the area. She requests that Sean to rescue Yosef Bryman, her best spy from the Germans who have captured him. After rescuing Yosef, Sean returns to Luc, to learn that Luc has planned a meeting with the resistance leaders.
However the meeting is ambushed by the Germans and the leaders and Sean are forced to flee. the Resistance set up a new HQ in the catacombs below Paris, there Luc reveals his plan to stage an operation at the Paris Grand Prix. Many high ranking Nazi officials and military officers will be attending the event and an Iron Cross will be presented to the winner, by a General Eckhardt. Sean, because of experience is forced by Luc to be the race car driver, but doubts that he can win the race without a competitive racing car. Sean's pre-war racing car has been found in the possession of none other than General Eckhardt. Sean infiltrates Eckhardt's villa and then proceeds to steal the car and escapes in it back to the Belle de Nuit.
Sean is sent by Wilcox to rescue, Dr. Kessler a German scientist and sneak him out of France. Sean manages to rescue Maria Kessler, to attempt to bring her father into working for the Allies, and return her safely to an HQ only to find the HQ under attack by the Nazis. After escaping with Skylar and the Kesslers family, Sean take them to the Catacombs, only to find that Veronique has become separated from them and was captured. In a fit of rage Sean, quits the resistance, after he finds out no one has a plan to rescue her.
Yosef helps Sean find Veronique who is,being held in the Palace of Justice close to Notre Dame.Sean using the information obtained by Yosef's connections helps Veronique and others escape. Sean and Veronique return to the Catacombs where they find out, since Sean quit the resistance Vittore had volunteered to drive in the Paris Grand Prix. However Sean proceeds to knock Vittore out and takes his place in the race knowing the car is wired with explosives. Sean asks Yosef before the race, to move the Kesslers' to the Belle De Nuit. Yosef before he dies warns Sean, by radio during the race, that the Belle de Nuit is under attack by Nazis. Sean wins the race and then drive the car into the box seats where the Nazi officers and officials are killed in the ensuing explosion, Sean escapes by ditching out the car at the last moment.
Sean returns to the Belle to find it has been ransacked, the Kessler Family gone and Vittore mortally wounded, who warns, Santos is a traitor and that the Nazis know about all the Resistance and are planning attacks. Sean returns to the catacombs and fights the Nazis, he finds Luc but Luc is caught in the collapsing rubble of a wall, Veronique shows mercy by killing Luc. Skylar contacts Sean requesting a meeting at the airport where she informs him that the Kesslers are being brought to the Doppelsieg factory,Veronique invites herself along.
Sean and Veronique go into the factory, with Skylar waiting at the plane. They soon reach the room where Sean and Jules were held, he breaks down the door to the room only to find Maria Kessler. Sean still sees the blood on the chair where Jules was executed. Sean asks Veronique to go with Maria, before blowing up the room and locating Dr Kessler. Dr Kessler explains that the factory is serving as cover for a concealed weapon which he has been forced to make. Sean manages to inflict critical damage and the facility begins to explode. Sean and Dr Kessler escape in the nick of time and meet up with Skylar,Veronique and Maria.
Skylar returns Veronique and Sean in Paris, where the whole city has risen in revolt against the German occupation. Sean and Veronique drive to the Eiffel tower where Dierker has established a temporary command post. The Resistance is openly fighting the Nazis in the streets, but Sean observes that they are weak without a leader and urges Veronique to take up the mantle leaving her to lead while he goes to the post. Inside the Eiffel Tower, Sean finds a drunken Dierker on the observation deck in the act of executing his own officers. Dierker and Sean exchange a few brief words, Dierker noting that is fitting that Sean is the one to kill him as he was the only one to survive. Sean shoots Dierker with a pistol, Dierker falls from the observation deck,to the ground. Veronique comes up and asks if it is all over, but Sean replies that he's just getting started.